I was unable to find anything on the 3d head movements for the lizards, and I don’t have the resources to deep dive into the code myself. sorry 
I was looking into the tiles also and found this page which is a cheat-sheet on tile editor and rendered tiles. I noticed that the tiles are possibly procedurally rendered shadows onto the outline image. I’m going to see if I can make a something like that where the shadows are drawn onto the outline of the tiles.
Also, if you’re wanting a creature to flip its segments based on where it’s facing, use this block:
It basically tells the sprites that have continuous rotation applied to flip vertically when changing direction. I created this block for my Chinese Dragon game.